| common name for an elongated, limbless reptile of the order Squamata, which also includes the lizards. Most snakes live on the ground, but some are burrowers, arboreal, or aquatic; one group is exclusively marine. In temperate climates they hibernate. They are generally solitary in their habits, although they may congregate in places offering food or shelter, and large numbers may hibernate together. Snakes range in length from about 4 in. (10 cm) to over 30 ft (9 m). Most are protectively colored. Characteristics Anatomy Snakes constitute the suborder Serpentes (or Ophidia). In most snakes limbs are entirely lacking, but a few have traces of hind limbs. The skin, which is covered with horny scales, is shed, usually several times a year. The extremely long, narrow body is associated with distinctive internal features. The number of vertebrae is much larger than in most vertebrates, paired internal organs are arranged linearly rather than side by side, and only one lung is developed, except in members of the boa family, which have two lungs. The jaws of snakes are loosely jointed and extremely flexible. The pointed, backward-curved teeth are fused to the supporting bones of the head. There are no ears or movable eyelids. Snakes have good vision. They do not hear airborne sound waves, but can perceive low-frequency vibrations (100–700 Hz) transmitted from the ground to the bones of the skull. A chemosensory organ opens into the roof of the mouth; it receives stimuli from the forked tongue that constantly tastes the surroundings as the animal moves along. Snakes have no larynx or vocal chords, but are capable of producing a hissing sound. Locomotion and Limblessness A snake moves by means of muscular contraction, which can produce several types of locomotion, the commonest types being undulation and straight-line movement. Straight-line movement is aided by the ventral plates, elongated scales on the abdomen that overlap with their open ends pointing toward the tail. These plates can be moved forward by means of muscles attached to the ribs. It is believed that snakes are descended from lizards, but how and why they evolved toward limblessness is uncertain. Some paleontologists have held that limblessness was an evolutionary advantage in the dense vegetation that formed the early environment of snakes, or that it developed to facilitate burrowing habits, but others believe that the earliest snakes evolved in a marine environment and are descended from marine lizards. Support for the latter view comes from recent discovery of the most primitive snake now known, a fossil specimen with two short but well-developed hind legs found in marine sediments in Israel. It lived in a shallow sea 95 million years ago. Predation Small snakes feed on insects and larger ones on proportionately larger animals. Their teeth are designed for catching and holding prey, but not for chewing. The construction of the jaws, the ribs, and the expandable skin enable them to swallow very large prey whole. Some snakes capture animals by pinning them to the ground; some—the constrictors—crush them by wrapping their bodies around them and squeezing; still others—the venomous snakes—inject poison into their victims. The poison, or venom, is produced by modified salivary glands from which it passes through either a groove or a hollow bore in the fangs, the enlarged, specialized teeth found in venomous snakes. A snake may bite a person when threatened or alarmed; if the snake is venomous the bite can sometimes prove fatal (see snakebite). Only by familiarity with the appearance of particular species, or by examination of the fangs, can the venomous snakes be distinguished from the harmless ones. Reproduction Fertilization is internal in snakes; as in lizards, the males have paired copulatory organs, either of which may be used in mating. Females of some species can store sperm for several years to insure future fertilization. In most species the female lays eggs; in some the eggs are incubated and hatched within the mother's body; in a few there is true viviparity, or live birth, with the young nourished by means of a placenta rather than an egg. Some egg-laying snakes brood the eggs, but there is no parental care of the young. Types of Snakes The approximately 2,700 snake species, of which about four fifths are nonvenomous, are distributed throughout the temperate and tropical zones of the world (except in New Zealand, Ireland, and some isolated oceanic islands) and are found in greatest profusion in the tropics. About two thirds of all snake species belong to the family Colubridae; most of these are nonvenomous. Among the harmless colubrid snakes of North America are the garter snakes (including the ribbon snake), the water snakes, the green, or grass, snakes, the black snakes, the racers, the king snakes (including the milk snake), and the bull, hognose, and rat snakes. The family Boidae (boas and pythons) includes the world's largest snakes, the South American anaconda and the Asian reticulated python, as well as the smaller boa constrictor and the tree and sand boas. Most poisonous New World snakes belong to the pit viper family; these include the copperhead, water moccasin, rattlesnake, fer-de-lance, and bushmaster. Venomous Old World snakes are the true vipers, including the adder and the asp, and members of the cobra family, including the mamba of Africa and the krait of Asia. The poisonous coral snakes of the New World also belong to this family. The venomous sea snakes inhabit tropical oceans. Importance Snakes are of major importance as pest controllers because of their extensive predation on destructive mammals such as rats and mice. Some, like the sea snakes and pythons, are highly regarded as food in Asia but, although most are probably edible, snakes are not widely used for meat. The skin is often used for belts, bags, and shoes. Venom is removed from snakes for use in treating certain diseases and to make antivenin for snakebites. See also snake worship. Classification Snakes are classified in the phylum Chordata, subphylum Vertebrata, class Reptilia, order Squamata, suborder Serpentes. Bibliography See A. H. and A. A. Wright, A Handbook of Snakes of the United States and Canada (2 vol., 1957); K. L. Williams and V. Wallach, Snakes of the World (2 vol., 1990). ____________________The Columbia Encyclopedia, Sixth Edition Copyright© 2004, Columbia University Press. Licensed from Lernout & Hauspie Speech Products N.V. All rights reserved. -44323- | 
