1809–82, English naturalist, b. Shrewsbury; grandson of Erasmus Darwin and of Josiah Wedgwood. He firmly established the theory of organic evolution known as Darwinism. He studied medicine at Edinburgh and for the ministry at Cambridge but lost interest in both professions during the training. His interest in natural history led to his friendship with the botanist J. S. Henslow ; through him came the opportunity to make a five-year cruise (1831–36) as official naturalist aboard the Beagle. This started Darwin on a career of accumulating and assimilating data that resulted in the formulation of his concept of evolution and his explication of natural and sexual selection. He spent the remainder of his life carefully and methodically working over the information from his copious notes and from every other available source.

Independently, the naturalist A. R. Wallace had worked out a concept of evolution similar to Darwin's. Wallace sent a paper outlining his theory to Darwin in 1858, and its striking coincidences with Darwin's work led Darwin's friends to move to assure that the more cautious Darwin, who had been slow to publish, would receive credit for the independence and priority of his ideas. The next year Darwin set forth the structure of his theory and massive support for it in the superbly organized On the Origin of Species, supplemented and elaborated in his many later books, notably The Descent of Man (1871). He also formulated a theory of the origin of coral reefs.

See his autobiography (ed. by N. Barlow, 1958) and Life and Letters (ed. by F. Darwin, 1887; repr. with intro. by G. G. Simpson, 1962); letters of Darwin and Henslow, ed. by N. Barlow (1967); The Corespondence of Charles Darwin ed. by F. Burkhardt et al. (10 vol., 1987–97); J. Barzun, Darwin, Marx, Wagner (rev. ed. 1958); G. Wichler, Charles Darwin: The Founder of the Theory of Evolution and Natural Selection (tr. 1961); A. Moorehead, Darwin and the Beagle (1969, rev. ed. 1979); P. Appleman, ed., Darwin (1970, repr. 1983); D. L Hull, Darwin and His Critics (1983); R. J. Richards, Darwin and the Emergence of Evolutionary Theories of Mind and Behavior (1989); R. Dawkins, River Out of Eden (1995); D. C. Dennett, Darwin's Dangerous Idea (1995); N. Eldredge, Reinventing Darwin (1995); S. Jones, Darwin's Ghost: "The Origin of Species" Updated (2000); J. Browne, Charles Darwin: Voyaging (1995) and Charles Darwin: The Power of Place (2002).

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