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cause it has a sensitive cortex which on
stimulation releases calcium to the interior.
This calcium then produces a reaction akin
to blood clotting, a reaction which is in-
hibited by dilute solutions of fat solvents.The theory that I have proposed has
much still to explain. If it is correct it will
serve as a guide for the study of the col-
loidal interpretation of protoplasmic be-
havior. But at best the theory is only a
bare outline, and there are huge gaps which
remain to be filled in. Each phase of the
theory requires amplification, and there
are dozens of unsolved problems which
stare us in the face. For example we are
constantly inquiring as to the mechanism
of calcium release from the cortex, and we
are also busily trying to interpret various
aspects of the clotting reaction, aspects
which I did not have time to discuss with
you. Moreover, we have hopes of relating
calcium release to the electric phenomena
which typically accompany excitation.
These are but a few of our problems. The
fact that the theory suggests these prob-
lems is a point in its favor. Right or
wrong, the theory deserves discussion and
test, for at the present time, in view of the
recent papers which have done much to
demolish the experimental support for the
permeability theory, there is really no other
theory to tie to.
REFERENCES CITED
ANGERER, C. A. 1936. J. Cellular Comp. Physiol.,
8: 329.
-----. 1937. Anat. Rec., 70 Supp.: 52.
CHURNEY and KLEIN. 1937. Biol. Bull., 72: 384.
DAHLGREN. 1915. Carnegie Inst. Wash., Pub. No.
212: 213.
DAUGHERTY. 1937. Physiol. Zool., 10: 473.
HEILBRUNN, L. V. 1928; 1937. The Colloid Chem-
istry of Protoplasm. Berlin; An Outline of
General Physiology. Philadelphia.
-----. 1934. Biol. Bull., 66: 264.
-----. Biol. Bull., 71: 299.
HEILBRUNN, L. V. and DAUGHERTY. 1938. Physiol.
Zool., 11: 383.
-----, -----. 1939. Physiol. Zool., 12: 1.
MAZIA and CLARK. 1936. Biol. Bull., 71: 306.
MOSER. 1939. J. Expt. Zool., 80: 423.

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Publication Information: Book Title: The Cell and Protoplasm. Contributors: Forest Ray Moulton - editor. Publisher: Science Press. Place of Publication: Washington, DC. Publication Year: 1940. Page Number: 198.
    
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