the cohorts and maniples. That is, they rushed for safety into the intervals between the cohorts (in signa), and also even into the smaller intervals between the maniples (in manipulos). Here signa seems to refer to the cohorts.
§ 14. The musical instruments used in the Roman army were the bugle (buccina, Fig. 5), the trumpet (tuba, Fig. 6), the cavalry trumpet (lituus, Fig. 7), and the horn (cornu). This last was made of the horn of a buffalo, and provided with a silver mouthpiece. The others were probably of brass.
B. G. II, 201 VII, 472. C. II, 35.
The Romans knew very well a fact familiar to modem tactics, that, to carry a command amid the tumult of battle or down a long line of march, the penetrating notes of a brazen horn are much more effective than the sound of the human voice. And accordingly the signals for the various evolutions of march and battle were given by horn and trumpet, first by the horn, at command of the general, then taken up by the trumpets. The bugle seems to have sounded the divisions of the day—reveille, noon, and night-fall.